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Esther Smith as Nikki Newman, an employee at a car rental company who wants to become a mother with partner Jason. They later become parents to Princess and Tyler.
Rafe Spall as Jason Ross, a former English teacher turned taxi driver who wants to become a father with partner Nikki. They later become parents to Princess and Tyler.
Sian Brooke as Karen, Nikki's sister, Scott's wife and Stevie's mother. She is initially a teacher but becomes a novelist.
Darren Boyd as Scott, Karen's eccentric husband and father to Stevie.
Ophelia Lovibond as Erica (series 1–2), Nikki's best friend and Freddy's ex-wife.
Oliver Chris as Freddy, Jason's best friend who owns a sporting goods company.
Imelda Staunton as Penny (series 1–2), Nikki and Jason's adoption officer.
Phil Davis as Vic Ross, Jason's father.
Paula Wilcox as Sandra Ross (series 1–3), Jason's mother.
Marian McLoughlin as Jilly Newman, Nikki's mother.
Roderick Smith as John Newman, Nikki's father.
Robyn Cara as Jen (series 1–3), Nikki's younger co-worker and friend who is quirky and learning to be an adult.
Fola Evans-Akingbola as Harper (series 2), Freddy's girlfriend after his divorce.
Scarlett Rayner as Princess (series 4–5), Nikki and Jason's daughter and Tyler's sister, age 16
Eden Togwell as Princess (series 2–3), age 10
Cooper Turner as Tyler (series 4–5), Nikki and Jason's son and Princess' brother, age 12
Mickey McAnulty as Tyler (series 2–3), age 6
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